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Endogenous production of NO occurs primarily through the enzymatic conversion of L-arginine (L-Arg) by nitric oxide synthase (NOS) [1, 2], which belongs to a family of enzymes that includes neuronal nitric oxide synthase (nNOS), inducible nitric oxide synthase (iNOS), and endothelial nitric oxide synthase (eNOS) [3]

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The first step is mediated by -glutamylcysteine synthetase (GCLC), which catalyzes the -amide bond between the -carboxyl group of glutamate and the amino group of cysteine to form -glutamyl-cysteine (-Glu-Cys)